When the Indo-Iranian peoples descended out of the Steppe into the Iranian plateau in the second millennium BCE, they did not enter empty land. They entered a country that had been settled, cultivated, and ritually organized for the better part of ten thousand years. The religion that emerged from the encounter — what would eventually become Zoroastrianism — was not Aryan religion in pure form. It was Aryan ritual technology running on a Neolithic Iranian substrate.

The previous four articles in this series have traced what the Indo-Iranian peoples carried with them: the linguistic inheritance documented by the Gathas and the Rigveda, the institutional architecture formalized at the BMAC, the pastoral religion of the Steppe, and the shamanic substrate that fed into all of it. Each layer was older than the one above it, and each layer descended ultimately from somewhere north of the Iranian plateau itself.

This article reverses the direction. The Iranian plateau, when the Aryans arrived, was already one of the oldest continuously inhabited religious landscapes in the world. The Zagros mountains and the high plains east of them had supported settled communities since at least the tenth millennium BCE — communities with their own gods, their own ritual architecture, their own religious specialists, and their own theological vocabulary. We can barely read this tradition directly, because the people who carried it left no surviving texts of their own. But we can recover enough of it to know two things with confidence: first, that it was substantial and ancient; and second, that when it met the Aryan religion descending from the north, the resulting synthesis — running on what the user's original framing called the Zagros hardware — was the actual material from which the historical religion of Iran was made.

I. Ten Thousand Years on the Plateau

The Zagros mountain range, running roughly north-west to south-east from eastern Turkey down to the Strait of Hormuz, is one of the four or five places on Earth where agriculture was independently invented. Between approximately 9500 and 8000 BCE — the period archaeologists call the Pre-Pottery Neolithic — communities along the western flanks of the Zagros and out onto the adjacent Iranian plateau began the domestication of goats, sheep, wheat, and barley. These communities, at sites such as Ganj Dareh in western Iran, Tepe Asiab, and Sheikh-e Abad, are among the earliest farming villages in the world.

The continuity that follows is the single most important fact about the Iranian plateau. From Ganj Dareh in the ninth millennium BCE to Tepe Sialk founded around 6000 BCE to Susa founded around 4200 BCE to the Achaemenid capitals of Pasargadae and Persepolis — there is no break in the human settlement of the plateau. Other regions saw demographic collapses, cultural ruptures, replacement migrations. The Iranian plateau, throughout this entire span, was continuously occupied by populations who buried their dead in the same places, worshipped on the same hills, and elaborated their religious practice as one continuous tradition for the better part of ten thousand years.

By the time the Indo-Iranian-speaking peoples began arriving in the plateau in significant numbers — toward the end of the second millennium BCE — that tradition had already developed for eight thousand years. The Aryans did not bring religion to a religionless country. They brought a particular ritual technology and a particular linguistic and theological vocabulary into a land that already had its own.

II. The Genetic Hardware

The continuity of the plateau population is not only an archaeological inference. It is now documented at the level of DNA. The 2016 ancient genomics study led by Iosif Lazaridis, sampling Neolithic individuals from Ganj Dareh and other Zagros sites, identified a distinct ancestral genetic component which has come to be designated Iran_N — the Iranian Neolithic ancestry. This component, the study found, was already present on the plateau in the early ninth millennium BCE and has persisted in the regional gene pool from that point to the present.

The Iran_N component is the dominant ancestry signal across a vast arc of populations. It constitutes roughly 60 percent of modern Iranian genetic ancestry, with the remainder accounted for by Steppe-derived (Indo-Iranian) and additional Near Eastern components. It contributes at least 50 percent — by some estimates more — of the ancestry of contemporary Levantine populations, with the remainder being Natufian/Semitic-associated ancestry. The Iran_N signal also contributed substantially to the populations that moved east into the Indus Valley and that contribute today to South Asian ancestry.

The genetic record makes the cultural argument visible at biological resolution. The same population that built the Neolithic villages of the Zagros, that founded Susa, that elaborated Elamite civilization across three millennia, and that absorbed the Indo-Iranian newcomers in the late second millennium BCE — is the same population that constitutes the majority of modern Iranian ancestry today. The continuity is not metaphorical. It is genetic.

Hardware and Software

The framing this series began with — Aryan religious culture as the software that ran on Zagros DNA as the hardware — is now confirmed at the level of population genetics. The dominant component of modern Iranian ancestry is the Iranian Neolithic substrate, not the Steppe input. The Indo-Iranian newcomers contributed language, ritual vocabulary, and a particular institutional theology. The continuous population of the plateau contributed the actual bodies, the agricultural economy, the urban infrastructure, and — as this article will argue — a substantial portion of what the resulting synthesis became.

III. Göbekli Tepe and the Neolithic Religious Revolution

To understand what kind of religion was already operating on the Iranian plateau when the Aryans arrived, the broader Near Eastern Neolithic must be set in view. Just to the west of the Zagros, in what is now southeastern Turkey, the site of Göbekli Tepe — excavated by Klaus Schmidt from 1995 until his death in 2014 — has rewritten the assumptions of Neolithic archaeology more thoroughly than any other find of the last half-century.

Göbekli Tepe is a monumental ritual complex dated to approximately 9600 to 8200 BCE. It consists of multiple stone enclosures built around T-shaped limestone pillars up to five meters tall, many of them carved with reliefs of animals — boars, foxes, scorpions, vultures, snakes, lions. The complex was built by hunter-gatherers before the regional transition to settled agriculture, not after. This is the single most consequential fact about the site. The conventional story had complex religious architecture emerging as a consequence of agricultural surplus. Göbekli Tepe inverts the order: monumental ritual architecture appears first, and the agricultural transition follows it.

The French archaeologist Jacques Cauvin had argued for this inversion on theoretical grounds in his 1994 book The Birth of the Gods and the Origins of Agriculture, published in English translation in 2000. Cauvin's argument — that a Neolithic religious transformation preceded and motivated the agricultural transition rather than following from it — was speculative when he made it. Göbekli Tepe is its archaeological vindication. The religious revolution came first.

Göbekli Tepe is not on the Iranian plateau. But it is the closest contemporary parallel for what the Iranian Neolithic was developing in the same centuries, and it establishes the basic point: the populations that became the Iran_N ancestry component were not simple early farmers with rudimentary religion. They were participants in a regional Neolithic religious culture whose sophistication is now archaeologically attested.

IV. The Plateau Sequence

The Iranian plateau itself has not produced a Göbekli Tepe — or has not yet, since less than one percent of likely sites have been systematically excavated. What it has produced is a continuous sequence of progressively more elaborate settlements, each preserving traces of the religious tradition the next layer built upon.

Ganj Dareh and the Early Neolithic (c. 8000 BCE)

At Ganj Dareh in the Kermanshah region, excavated by Philip Smith from 1965 to 1973, the earliest Iranian Neolithic village preserves the foundations of a religious life that would prove extraordinarily persistent. The site contained clay figurines, including animal figures and stylized human forms, evidence of a developed iconographic tradition. Burials were placed beneath the floors of houses — a practice with deep ritual significance, documented across the Near Eastern Neolithic, and indicating an integration of the dead into the continuing life of the household that would persist in Iranian religious sensibility for millennia.

Çatalhöyük and the Wider Context (c. 7400 BCE)

Çatalhöyük, in central Anatolia, lies outside the strict Iranian plateau but represents the same Neolithic religious world in which the Zagros Neolithic was embedded. Excavated extensively by James Mellaart and then by Ian Hodder, Çatalhöyük preserved an extraordinary repertoire of religious imagery: bull horns mounted in domestic shrines, leopard reliefs, mother-figure statuettes, vulture imagery associated with excarnation rituals, and again the practice of sub-floor burial. The bull, in particular, would prove to have an extremely long life in the religious imagination of the broader region — eventually reappearing in Achaemenid imperial iconography on the bull-capitals of Persepolis.

Tepe Sialk and the Rise of Monumental Ritual (c. 6000–3000 BCE)

Tepe Sialk, in what is now the Kashan region of central Iran, is perhaps the most important site for understanding the continuity from the Zagros Neolithic to the historical Iranian world. Founded around 6000 BCE and continuously occupied for thousands of years, Sialk eventually produced — by approximately 3500 BCE — a tall mudbrick platform whose form anticipates by two millennia the later Mesopotamian ziggurat and the still later Achaemenid ritual platform. The Tepe Sialk platform is among the earliest examples of monumental religious architecture on the Iranian plateau and establishes the platform-temple form as a deep continuity in the regional tradition.

Susa and the Elamite Continuum (c. 4200 BCE – 550 BCE)

Susa, founded around 4200 BCE in what is now southwestern Iran, became the principal city of the Elamite civilization that would dominate the western plateau through the entire third and second millennia BCE and into the first. The Elamites are crucial to the argument of this article. They were the historical successors of the Zagros Neolithic tradition. Their religion — known partly from their own surviving inscriptions and partly from Sumerian and Akkadian references — preserved a recognizably ancient and distinct theological vocabulary, organized around such figures as Inshushinak (the patron of Susa), Humban (a sovereign deity), and Kiririsha (a mother-goddess of broad cosmic function).

The Achaemenid Persian Empire that emerged in the sixth century BCE did not erase Elamite tradition. It absorbed it. Elamite was one of the three official languages of the Achaemenid administration, alongside Old Persian and Akkadian. Elamite scribes drafted the imperial documents. Elamite ritual personnel served at the Persian courts. The Persepolis Fortification Archive, the principal documentary witness to Achaemenid administration, is written almost entirely in Elamite. The substrate did not disappear under the Aryan reform. It became the institutional backbone on which the Aryan reform built its empire.

V. What the Aryans Encountered

The convergence of these strands of evidence allows a clearer picture of the religious situation on the Iranian plateau at the moment of Indo-Iranian arrival — sometime between approximately 1500 and 1000 BCE, depending on the region. By that point, the plateau was already home to:

A continuously occupied agricultural population in continuity with the Neolithic populations of eight thousand years earlier, carrying the Iran_N genetic component that would constitute the demographic majority of every subsequent population on the plateau.

An established urban civilization in the south (Elam, centered on Susa and Anshan) with a developed religious vocabulary, monumental temple architecture, a literate priesthood writing in Linear Elamite and later in Elamite cuneiform, and a continuous theological tradition reaching back through Proto-Elamite into the regional Neolithic.

A regional iconographic tradition — bull-cult, mother-figure, water-goddess, mountain-sanctuary — that had been developing across the broader Near Eastern Neolithic for millennia and that was already present in the plateau material from at least the seventh millennium BCE.

A platform-temple architectural tradition attested at Tepe Sialk by 3500 BCE, predating the comparable Mesopotamian ziggurat tradition and establishing the raised ritual platform as a regional architectural type.

And, almost certainly though now invisible to direct documentation, a religious specialist class whose office and function the arriving Aryan priesthood would in some measure absorb. The Indo-Iranian title athravan (priest) descends from the Steppe inheritance. But the institution of the priesthood, the daily ritual cycle, and the urban temple complex were already present on the plateau before the Aryans arrived. The Aryan priestly office did not need to invent these structures. It needed only to occupy them.

VI. The Hardware that Carried the Fire

What survived of the Zagros substrate in the resulting synthesis can be traced — partially — in the features of the historical Iranian religion that have no clear Indo-Iranian or Steppe parallel and that do correspond to identifiable plateau-Neolithic precedents.

Substrate Element Neolithic Plateau Evidence Survival in Historical Iranian Tradition
The bull as sacred animal Çatalhöyük bull horns; Susa bull iconography; the bull on Iranian seals The bull-capitals at Persepolis; the bull-slaying motif preserved in Mithraism; Gavaevodata as the primordial ox of Zoroastrian cosmology
The raised ritual platform The Tepe Sialk platform (c. 3500 BCE); the Elamite ziggurat at Chogha Zanbil (c. 1250 BCE) The Achaemenid apadana on its stone platform; the raised plinth of the Zoroastrian fire-temple
The mother / water goddess Female figurines from Ganj Dareh, Tepe Sarab, and across the Iranian Neolithic; Elamite Kiririsha Anahita — the water-goddess absorbed into Achaemenid imperial worship and recognized in the Avesta; one of only two Iranian deities to receive monumental cult statues
Sub-floor burial & the integration of ancestors Documented at Ganj Dareh, Çatalhöyük, and across the regional Neolithic The Achaemenid royal tombs at Naqsh-e Rustam carved into the living rock; the persistent Iranian sensibility of the dead as continuing presence in the landscape
The mountain sanctuary Open-air ritual sites on plateau highlands attested from the Neolithic onward The Zoroastrian preference for elevated and exposed sites; the Mithraic mountain associations; the sanctity of named peaks across the Iranian highlands
The literate priesthood & temple economy Elamite temple administration documented from the third millennium BCE The Achaemenid administrative bureaucracy at Persepolis (Elamite cuneiform); the Sasanian temple-estate system; the role of the mobed as administrative as well as ritual figure

None of these continuities, taken alone, would settle the case. The bull is widely sacred. Platforms are built by many cultures. Goddess figurines are not unique to any single tradition. But the convergence of all six elements on the Iranian plateau, in a context of unbroken demographic continuity from the Neolithic to the present, makes the synthesis argument difficult to avoid. What the Aryans brought into the plateau encountered a tradition that was very nearly its equal in antiquity, and that was institutionally and economically more developed at the moment of contact. The encounter did not produce Aryan religion in pure form. It produced what we now call Iranian religion — a synthesis whose components can still be partially disentangled, but whose effective unity is itself the historical fact.

"The land was already old when the Aryans came down from the high places. The fire on the hilltops was lit by hands they had never known."

— Reconstructed framing, after Boyce 1975

Mary Boyce, in her foundational History of Zoroastrianism, was already careful to distinguish the inherited Indo-Iranian elements of the religion from the elements that the Iranian tradition acquired on the plateau itself. Her treatment of Anahita in particular — a deity not securely reconstructable to the Proto-Indo-Iranian level, whose iconography is consistent with the regional Neolithic mother-goddess tradition, and who is given monumental cult by Artaxerxes II in the fourth century BCE — is among the clearest cases. Anahita is the substrate visible at the surface. She is the Iran_N component speaking through the Aryan ritual.

The fire that Zoroaster faced when he prayed was a fire that the Aryans had carried out of the Steppe and the BMAC. But it burned on a stone platform whose architectural genealogy descended from Tepe Sialk. It was tended by priests whose institutional office had been functioning at Susa for two thousand years before Zoroaster was born. And it was understood by the population that received it through a religious sensibility shaped by ten thousand years of continuous Neolithic and Bronze Age tradition on the plateau.

This is the hardware. The five layers traced through Articles I–IV — Proto-Indo-Iranian liturgy, BMAC institutional architecture, Steppe pastoral religion, shamanic substrate, and now Zagros Neolithic substrate — are now all on the table. The final article of this cluster will trace how they converged.

Primary Sources & Scholarly Literature

1 Iosif Lazaridis, Dani Nadel, Gary Rollefson, et al., "Genomic insights into the origin of farming in the ancient Near East," Nature 536 (2016): 419–424. The principal study identifying the Iranian Neolithic (Iran_N) ancestry component from Ganj Dareh and demonstrating its persistence across subsequent populations.
2 Klaus Schmidt, Göbekli Tepe: A Stone Age Sanctuary in South-Eastern Anatolia (Berlin: ex oriente, 2012). The principal published treatment of the Göbekli Tepe excavations by the lead excavator; essential for understanding the Neolithic religious revolution in the broader region.
3 Ian Hodder, The Leopard's Tale: Revealing the Mysteries of Çatalhöyük (London: Thames & Hudson, 2006). The standard accessible account of the Çatalhöyük material; particularly valuable for the bull, leopard, and goddess iconography that connects to subsequent Iranian-plateau traditions.
4 Jacques Cauvin, The Birth of the Gods and the Origins of Agriculture, trans. Trevor Watkins (Cambridge: Cambridge University Press, 2000 [French original 1994]). The foundational theoretical argument that Neolithic religious transformation preceded the agricultural transition; subsequently vindicated by the Göbekli Tepe excavations.
5 Frank Hole (ed.), The Archaeology of Western Iran: Settlement and Society from Prehistory to the Islamic Conquest (Washington: Smithsonian Institution Press, 1987). The standard English-language reference work on the long Iranian-plateau sequence; still the indispensable starting point for the regional archaeology.
6 Hassan Fazeli Nashli and Roger Matthews (eds.), The Neolithisation of Iran: The Formation of New Societies, BANEA Publication Series 3 (Oxford: Oxbow Books, 2013). The current synthesis of Iranian Neolithic research; brings together two generations of plateau archaeology.
7 Daniel T. Potts, The Archaeology of Elam: Formation and Transformation of an Ancient Iranian State, 2nd ed. (Cambridge: Cambridge University Press, 2016). The standard treatment of the Elamite civilization; the principal documentation for the continuity from the Neolithic substrate through the Bronze Age into the Achaemenid period.
8 Pierre Amiet, L'âge des échanges inter-iraniens, 3500–1700 avant J.-C. (Paris: Éditions de la Réunion des Musées Nationaux, 1986). The most detailed treatment of the Iranian plateau's role as a cultural connector during the formative third and second millennia, including its contacts with Mesopotamia and the Indus Valley.
9 Ofer Bar-Yosef, "The Natufian Culture in the Levant, Threshold to the Origins of Agriculture," Evolutionary Anthropology 6 (1998): 159–177. Essential context for the broader Near Eastern Neolithic transition in which the Zagros sequence is embedded.
10 Trevor Watkins, "New Light on Neolithic Revolution in South-west Asia," Antiquity 84 (2010): 621–634. Synthesis of the cognitive, social, and religious dimensions of the Neolithic transition in the broader region, including the Iranian plateau.
11 C.C. Lamberg-Karlovsky and Maurizio Tosi, "Shahr-i Sokhta and Tepe Yahya: Tracks on the Earliest History of the Iranian Plateau," East and West 23 (1973): 21–57. Foundational treatment of the Bronze Age Iranian plateau as a connective cultural zone between the Indus Valley, the Persian Gulf, and Mesopotamia.
12 Mary Boyce, A History of Zoroastrianism, Vol. I: The Early Period (Leiden: Brill, 1975). Boyce's careful disentangling of Indo-Iranian inheritance from plateau-substrate elements in early Iranian religion; her treatment of Anahita is particularly important for the case that the Neolithic substrate is partially recoverable through the historical record.